In Ezekiel 26:11, this signifies the love of learning or intellectual things. (Arcana Coelestia 3727)

In Revelation 6:2, this signifies the love of understanding the Word. (Apocalypse Revealed 298)

In Zechariah 12:4, this signifies that the intellectual should be filled with falsities. (Arcana Coelestia 2383[2])

In general, 'a horse' signifies knowledge or understanding of the Word. In an opposite sense it signifies the understanding of the Word falsified by reasonings, and likewise destroyed from self-derived intelligence. 'A dead horse' signifies no understanding of truth from the Word.

In Genesis 1:24, beasts signify the things of man's will or loves. (Arcana Coelestia 44, 46)

In Genesis 9:10, beasts signify all that was living in the man of the Ancient Church, and also what belonged to his new will; likewise the lower things of his understanding and the will therefrom. (Arcana Coelestia 1026-1029)

In Psalm 104:20, beasts signify affections longing to be instructed, or spiritually nourished. (Apocalypse Explained 650[10])

In Luke 10:35, since the beast was a donkey, this signifies to instruct another according to his capability. (Apocalypse Explained 1154)

The beast of the south (Isaiah 30:6) signifies people who are principled in the knowledges of good and of truth, but do not apply them to life and instead to science.

Every beast and creeping thing (Genesis 8:19) signifies the goodnesses of the internal and external man.

"Beasts" represent the affection for doing good things, a true desire to do them from the heart. In the negative sense, "beasts" stand for the lust to do evil.

The beast ascending out of the sea (Revelation 13:1) signifies reasonings from the natural man confirming the separation of faith from life.

(Референце: Apocalypse Explained 13, 773; Revelation 13:11)