In Exodus 17:10, they signify divine truth in successive order, that is, divine truth immediately from the Lord, mediately from Him, and divine truth proceeding mediately from this. (Arcana Coelestia 8602)

9524. 'Which are over the ark of the Testimony' means with the Lord in heaven. This is clear from the representation of 'the ark' as heaven, and from the meaning of 'the Testimony' as Divine Truth, thus the Lord in heaven, both dealt with in 9503.